Tarantulas comprise a group of large and often hairy spiders of the family Theraphosidae. As of December 2023, 1,100 species have been identified, with 166 genera. The term "tarantula" is usually used to describe members of the family Theraphosidae, although many other members of the same infraorder (Mygalomorphae) are commonly referred to as "tarantulas" or "false tarantulas". Some of the more common species have become popular in the exotic pet trade. Many New World species kept as pets have setae known as urticating hairs that can cause irritation to the skin, and in extreme cases, cause damage to the eyes.
Like all arthropods, the tarantula is an invertebrate that relies on an exoskeleton for muscular support. Like other Arachnida, a tarantula's body comprises two main parts, the prosoma (or cephalothorax) and the opisthosoma (or abdomen). The prosoma and opisthosoma are connected by the pedicel, or pregenital somite. This waist-like connecting piece is actually part of the prosoma and gives the opisthosoma a wide range of motion relative to the prosoma.
Depending on the species, the body length of tarantulas ranges from about 5 to 11 cm (2 to 4+1⁄2 in) with leg spans of 8–30 cm (3–12 in).[citation needed] Leg span is determined by measuring from the tip of the back leg to the tip of the front leg on the opposite side. Some of the largest species of tarantula may weigh over 85 g (3 oz); the largest of all, the goliath birdeater (Theraphosa blondi) from Venezuela and Brazil, has been reported to attain a weight of 170 g (6 oz) and a leg-span up to 30 cm (12 in), males being longer and females greater in girth. The fang size of this tarantula reaches a maximum of 4 cm (1+1⁄2 in).
Theraphosa apophysis (the pinkfoot goliath) was described 187 years after the goliath birdeater, so its characteristics are not as well attested. T. blondi is generally thought to be the heaviest tarantula, and T. apophysis has the greatest leg span. Two other species, Lasiodora parahybana (the Brazilian salmon birdeater) and Lasiodora klugi, rival the size of the two goliath spiders.
Most species of North American tarantulas are brown. Elsewhere, species have been found that variously display cobalt blue (Cyriopagopus lividus), black with white stripes (Aphonopelma seemanni), yellow leg markings (Eupalaestrus campestratus), metallic blue legs with vibrant orange abdomen and green prosoma (Chromatopelma cyaneopubescens). Their natural habitats include savanna, grassland such as in the pampas, rainforest, desert, scrubland, mountains, and cloud forest. They are generally classed among the terrestrial types. They are burrowers that live in the ground.
Tarantulas are becoming increasingly popular as pets and some species are readily available in captivity.
Tarantulas can be confused with other members of the order Mygalomorphae, such as trapdoor spiders, funnel-web spiders and purseweb spiders. They can also be confused with some members of the order Araneomorphae such as the Lycosidae family. There are multiple ways to identify a tarantula. First the hairs: in the Americas most tarantulas have urticating hairs, though some, such as the Hemirrhagus genus, lack these. The hairs are usually more noticeable than with most other spiders. Another is the size, as tarantulas tend to be bigger, but this is again not a failproof way. They also do not use their webs for hunting, instead using them as building material or tripwire.
One of the most decisive ways to tell is by looking at their fangs. Tarantula fangs face downwards, as opposed to those of true spiders, which face each other, allowing them to make pincerlike motions. They also own two book lungs, as opposed to true spiders which only have one. Their lifespan is also longer than most spiders.
The spider originally bearing the name tarantula was Lycosa tarantula, a species of wolf spider native to Mediterranean Europe. The name is derived from the southern Italian town of Taranto. The term tarantula was subsequently applied to almost any large, unfamiliar species of ground-dwelling spider, in particular to the Mygalomorphae and especially the New World Theraphosidae. Compared to tarantulas, wolf spiders are not particularly large or hairy, and so among English speakers in particular, usage eventually shifted in favour of the Theraphosidae, even though they are not closely related to wolf spiders at all, being in a different infraorder.
The name tarantula is also incorrectly applied to other large-bodied spiders, including the purseweb spiders or atypical tarantulas, the funnel-webs (Dipluridae and Hexathelidae), and the dwarf tarantulas. These spiders are related to tarantulas (all being mygalomorphs) but fall into different families from them. Huntsman spiders of the family Sparassidae have also been termed tarantulas because of their large size, when, in fact, they are not related. Instead, huntsman spiders belong to the infraorder Araneomorphae.